Wonderful Alexander, the kitten who is the biggest, stongest, and loudest in his family, thinks he is destined for wonderful things. No sooner has he set out to explore on his own than he is chased up a tree and stuck there. His rescuer, Jane, a black kitten with wings, leads him to her home, where Alexander meets the other Catwings.

Alexander soon learns how he can repay Jane, who has been so wonderful to him. He helps Jane confront her greatest fear.