Life on earth has changed forever as humans and their courageous robotic allies, the Autobots, must work together to protect the planet from the destructive forces of the evil Decepticons. At the headquarters of NEST (Networked Elements: Supporters and Transformers), tech sergeant Epps and captain Lennox both guard and assist cyberneticist Yoki Ishihara and the brooding Russian AI genius Petr Ivanov as they explore the differences between organics and bots. In the meantime, all around them, alliances fray, distrust grows, suspicions mount, and traitors come out of the shadows.

Optimus Prime, the powerful leader of the Autobots who is also part of NEST, is on the defense as battles flare up from Australia to Zimbabwe. But escalating Decepticon attacks will culminate in a final confrontation from which no one-man or machine-will emerge unscathed.